They use the same senses as we do and their feet and hands are similar to
ours, chimpanzees still have opposable toes and can grasp things with their feet.
A. except what
B. except for
C. except that
D. besides that
网友回答
A. except what
B. except for
(答案→)C. except that
D. besides that
解析：that从句作except的宾语。