The manager spoke highly of such as loyalty, courage and truthfulness shown by his employ

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(答案→)A. virtues
B. features
C. properties
D. characteristics
解析：句意，这位经理高度评价他的员工展现出来的一些美德，例如：忠诚、胆量、诚实。Virtues 美德 features特征 properties特性 characteristics 特征，根据题意，故选A。